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Page number:287 
Remarks (public):The species is very similar to the North American E. tinctorium (Ellis & Everh.) Ellis & Everh., but this species is triquetrous and up to 10 cm thick, and with flat to conical spines with fibrous, sterile dissepiments, darker than the rest of the spine (Gross 1964). 
Description type:Non-original description 
Description:Echinodontium tsugicola (Henn. & Shirai) Imazeki
Jap. J. Bot. 11:517, 1935: Hydnofomes tsugicola Henn. & Shirai, Bot. Jb. 28:268, 1900.
Basidiocarps perennial, pileate to effused-reflexed, solitary or 2-3 imbricate, flat, 1-8 cm long x 1-4 cm broad, 1-2 cm thick; pilear surface dark brown to black, zonate, rimose-rugose, margin brown, undulating, pubescent; hymenophore irpicoid, light buff, spines scattered to adjacent, conical, up to 1.5 cm long and 1-2 mm thick, inner trama brick orange, apices acute to blunt, fertile; context brick red, woody, zonate, 0.5-1 cm thick.
Hyphal system dimitic; generative hyphae with clamps, thin-walled, 3.5-4.5 µm wide, hyaline, smooth, regularly branched; skeletal hyphae orange to rusty brown, thick-walled to solid, 3.5-4.5 µm wide, rarely branched.
Cystidia subulate to fusiform, encrusted in the upper half, 30-40 x 6-10 µm, protruding up to 15 µm, at first thin-walled and hyaline, becoming thick-walled and orange brown.
Basidia clavate, 20-30 x 6-8 µm, with four sterigmata.
Basidiospores ellipsoid, thick-walled, hyaline, strongly amyloid, minutely echinulate, 6-8 x 4-6 µm.
Substrata. Known on Tsuga diversifolia, T. sieboldii, and Ables firma. Distribution. Only known from cold-temperate Japan (Nagano and Tochigi in Honshu).
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