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 Add this item to the list   Globifomes graveolens (Schw.) Murr. Fig. 146.
   
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Page number:307 
Remarks (public):Globifomes graveolens is macroscopically unique among North American polypores. However, its microscopic characters are so similar to those of Fomes fomentarius and F..fasciatus that it could logically be considered congeneric with them. All have the distinctive sclerids in their granular context. They are also alike in having a hard, gray crustose layer on the pileus surface. 
Description type:Non-original description 
Description:Globifomes graveolens (Schw.) Murr. Fig. 146.
Bull. Torrey Bot. Club 31:424, 1904. - Boletus graveolens Schw., Schr. Nat. Ges. Leipzig 1:97, 1822.
Basidiocarps apparently annual or developing new tubes or pilei for 2-3 seasons but persistent for longer periods, sessile, ungulate to columnar, with large numbers of small, imbricate petaloid pilei developing over the surface of a mass of core tissue, up to 15 cm wide, 20 cm long and 15 cm thick, upper surface of pilei at first dull brown, with a pale tan margin, minutely tomentose, radially rugose, eventually becoming gray to grayish-black, glabrous and hard and crustose, pore surface at first purplish gray, becoming dark grayish brown, the pores circular, 5-7 per mm, with thick, entire dissepiments, context in individual small pilei yellowish brown, fibrous, up to 5 mm thick, azonate, with a distinct, dark, crustose upper layer, context of central core with a granular or mottled appearance from interspersed areas of light and dark mycelium, this central core making up most of the basidiocarp, up to 18 x 15 cm, tube layers not stratified but some buried in context tissue at the base of small pilei, pale purplish brown, up to 2.5 mm thick, tubes whitish within.
Hyphal system trimitic, contextual generative hyphae difficult to find in older specimens, hyaline, thin-walled, with clamps, 3.5-5.5 µm in diam, contextual skeletal hyphae of fibrous context pale brownish in KOH, thick-walled, nonseptate, with rare branching, 5-11 µm in diam, binding hyphae especially abundant in granular context, hyaline to pale brown in KOH, thick-walled, nonseptate, much branched, 1-4 µm in diam, sclerids abundant in granular core, irregular in size and shape, many lobed or branched, thick-walled, pale to dark reddish brown in KOH, negative in Melzer's reagent, walls up to 10 µm thick.
Cystidia fusoid, hyaline, thin-walled, not projecting, 17-25 x 4.5-5.5 µm, with a basal clamp.
Basidia clavate, 4-sterigmate, 22-27 x 7-9 µm, with a basal clamp. Basidiospores cylindric, hyaline, smooth, negative in Melzer's reagent, 10-14 x 3-4 µm.
Type of rot. - White rot of the heartwood of living hardwoods, and, according to Overholts (1953), also fruiting on wounds in living trees. Cultural characteristics. - See Davidson et al. 1942, Nobles 1948, 1958, 1965, Stalpers 1978.
Sexuality. - Unknown.
Substrata. - Known only on hardwoods with most of the records from Quercus. Fruiting may continue on recently killed or fallen trees (3, 26, 69, 104, 153).
Distribution. Through most of the eastern, midwestern, and southeastern U.S. to east Texas. Not known from western North America or elsewhere in the world.
 
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