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Page number:57 
Description type:Non-original description 
Description:RAMARIA CAMPESTRIS (Yokoy. & Sag.) Petersen, stat. nov.
Basionym: Ramaria zippelii var. campestris Yokoyama & Sagara. 1973. Trans. Mycol. Soc. Japan 14: 302.
Type (holotype): TNS - Yufoin-machi, À‹ita pref., 6.ix.68, coll. by residents, Sagara no. 1502. [!]
Figures 8-11.
Fruitbodies up to 15 cm high, massive, branched in up to 5 ranks. Stipe 3-7 x 3.5-5 cm, tapering downward, solid, smooth but not glabrous, not beset with primordial branchlets, white or whitish when fresh, bruising to weak chocolate brown ("warm sepia"); flesh soft, somewhat woody, whitish, brunnescent ("fawn color", "light russet-vinaceous", "army brown"). Major branches very few, arising as a bifurcation of the stipe apex, 1-2.5 cm thick, concolorous with stipe below, with upper branches above; branches many, yellow-ocher when young ("maize yellow", "baryta yellow", "buff yellow", "antimony yellow"), deepening through intermediate shades ("ochraceous orange", "yellow ocher", "ochraceous tawny", "Mars yellow", "raw sienna", "Buckthorn brown", "Dresden brown") to olivaceous brown at maturity ("buffy olive", "dark olive buff", "grayish olive"); internodes short, diminishing abruptly; axils rounded; apices coarse, dentate to digitate, white or whitish in middle age, darker than branches at maturity ("deep olive buff", "dark olive buff"). Odor faint; taste bitter.
Hyphae of branch trama up to 16 um diam, weakly yellow in mass, clamped, somewhat thick-walled. Hymenium thickening; basidia up to 75 x 12-13 µm, broadly clavate, hyaline, clamped, (2)-3-4-sterigmate; sterigmata up to 8 um long, narrowly conical, spindly. Spores 11.5-14.1 x 5.6-6.7 um (E = 1.89-2.38; _Em = 2.07; Lm = 12.9 um), short-cylindrical to broadly ellipsoid, dull ocher to ochraceous tan in prints ("honey yellow", "yellow ocher", "clay color"); contents multiguttulate when young, finally uniguttulate at maturity; wall up to 0.5 um thick; hilar appendix gradual, broad, leaving a moderate "throat"; ornamentation of thorns up to 1.5 um long, sharp, tapering gradually, scattered over entire wall surface.
Macrochemical reactions: not reported.
Among grasses in meadows, forming fairy rings; Kuju Highlands, Kyushu, Japan.
OBSERVATIONS: The spores of this fungus seem similar to those of R. cokeri (= R. fragillima ss. Corner), and the basidia are basically 4-spored as wella Fruitbody stature and
color, however, are very different. As shown in the color illustrations by Yokoyama & Sagara, immature fruitbodies of R. campestris are very stout and stocky, and are yellow to yellow ocher in color. Those of R. cokeri are slenderer, and brick red. Moreover, R. cokeri is a forest inhabitor, white R. campestris seems to have a special (mycorrhizal?) relationship with bamboo grass (Pleioblastus distichus var. nezasa Mur. & Okam.). In fact, in stature, young fruitbodies seem to resemble young stages of R. grandis, but of very different color.
To my knowledge, R. campestris is the sole member of the subgenus to prefer a pastoral babit, but it is known only from its general topotype area. A specimen (13.ix.78, "on ground in Pinus densifolia - Quercus eerrata- forest", Yokoyama no. 2661) kindly furnished by. Dr. K. Yokoyama, indicates that its habitat may be more varied than hitherto reported.
The specimens cannot be accomodated by R. zippelii (ss. mihi), for that species exhibits characteristic 2-sterigmate, golden-refringent basidia (as does R. grandis, to which R. campestris bears more superficial resemblance). For this reason, it bas been proposed at the species rank.
 
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