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Page number:112 
Remarks (public):LLOYD recorded the Japanese collections under two names, Yasuda 98 as Hydnum imbricatum (1912, Lett. 42: 15), and Yasuda's unnumbered collection as Hydnum aspratum (1921: 1095), although he did not fail to observe that he saw little merit in their specific distinction.
The interesting point about Yasuda's collections is that the former - consisting of two young basidiomes - was said to have no odour, whereas the latter -a very old and weathered specimen, having lost most of its scales and spores-was stated to be fragrant.
This reintroduces the problem of the identity of Hydnum aspratum. As pointed out earlier (MAAs GEESTERANUS, 1960: 344), I regarded the type of that species as conspecific with S. imbricatus and, until I am shown to be wrong, I see no reason to change my mind. But I am now prepared to believe that there exists a second species which the Japanese continue to call Sarcodon aspratus, although it would seem to represent a different but closely allied species. This would differ from S. imbricatus in the narrower scales on the pileus and the all pervading smell. The latter character, however, does not seem to be entirely reliable, judging from Yasuda's error of mistaking a weathered S. imbricatus, with most of its characteristic features gone, for S. aspratus.
The consequence of the above is that the various Japanese records for "Sarcodon aspratus" have been left out of consideration, as they may refer to a species different from S. imbricatus.
In Lloyd's collection there are also, from China, a specimen and a fragment of the pileus of a second one, which were recorded (LLOYD, 1924:1320) under Hydnum subsquamosum. Both specimens represent S. imbricatus.
VASUDEVA (1962: 48) reported the present species from Mawphlong, Assam, adding that it had been found growing on coniferous wood. This sounds like some error, but since I fail to make out from what source the author had his information, I prefer to ignore the record.
The specimen in Lloyd 4904 used for investigation appeared to have an excess of undersized spores, frequently measuring only 6.5 µm in length or even less. It should be observed, however, that the spores in this weathered specimen remain only near the tips of the spines. It is possible that these spores are generally somewhat smaller than those of the earlier generations produced in the middle and nearer the base of the spines
 
Description type:Non-original description 
Description:SARCODON IMBRICATUS (L. ex Fr.) P. Karst. -Figs. 154-156
Basidiome simple. Pileus up to about 150 mm across, more or less regularly orbicular or lobed, plano-convex with depressed or umbilicate centre, eventually becoming infundibuliform and occasionally with a central hole which may extend to the base of the stipe; scaly from the beginning but losing its scales with advancing age; scales in concentric rings, coarse and thick and upright in the centre, imbricate and flat and decumbent farther outwards, passing into appressed fibrils near the margin, more or less dark brown on a yellow-brown ground. Stipe 50-120 x 7-20 mm, central to somewhat excentric, stocky to slender, equal or somewhat enlarged below, usually with attenuate base, fibrillose, glabrescent, solid at first but becoming hollow under certain conditions, whitish, becoming progressively brown with age, base covered with white mycelium. Spines up to about 9 mm long, decurrent, crowded, subulate, finally dark chocolate brown. Flesh brownish pallid in the pileus, somewhat darker in the stipe, neither blackish nor greenish in the base.
Context monomitic, consisting of generative hyphae. Hyphae up to 21.5 µm wide, inflating, thin-walled, branched, septate, with large clamp connections. Context of the spines similar, but hyphae narrower, up to 9 µm. Basidia 30-40 x 7-8 µm, clavate, with basal clamp, 4-spored, with sterigmata 3.6-5.4 µm long. Spores (7.2-)7.6-8.8 x 4.6-5.2 µm, of irregular outline, tubercular (tubercles numerous, strongly prominent, coarse, flat-topped to exsculpate), brownish, with oblique apiculus. Cystidia absent.
The above description is based mainly on the few dried collections available.
 
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