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 Add this item to the list   Climacodon septentrionalis
   
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Page number:65 
Remarks (internal):C. septentrionalis is morphologically variable. The pilei can be very thin, especially when they are reaching their maximal size. At the beginning of growth, when the pilei are thick and tuberculate, C. septentrionalis may be mistaken for Creolophus cirrhatus, but the structure of the context is quite different.
C. septentrionalis resembles the polypore Climacocystis borealis (Fr.) Kotl. & Pouz. surprisingly closely. The colour, gross morphology and consistency of the two species are similar, they are both very strongly fibrous inside, and the monomitic hyphal structure, cystidia (this similarity also noted by Domanski 1981) and other microscopic characters show a close resemblance. The cultures of both species produce chlamydospores and they give similar results in laccase and peroxidase tests (Stalpers 1978). C. borealis has often been regarded as a brown-rot fungus, but in fact it causes a white-rot (Stalpers 1978, Jülich 1984, Gilbertson & Ryvarden 1986), like C. septentrionalis. Bendiksen (1986) and Courfenay & Burdsall (1982) even listed C. borealis under the generic name Climacodon, but they were unintentional and invalid combinations (Bendiksen in litt., Burdsall in litt.), caused by the similarity of the two generic names. A closer comparison of Climacocystis and Climacodon would be interesting. The production of chlamydospores, type of cystidia,fibrous structure and hence somewhat fan-shaped pilei, and many other features also bring to mind the genus Oligoporus in the sense of Erkkilä and Niemelä (1986).
 
Description type:Non-original description 
Description:Climacodon septentrionalis
Hydnum septentrionale Fries, Syst. Mycol. 1: 414, 1821. - Climacodon septentrionalis (Fr.) Karst., Rev. Mycol. 3(9): 20, 1881. - Type locality: 'Suecia superior' (Fries 1821), meaning mountainous or otherwise elevated area of Sweden; Maas Geesteranus' (1974: 566) interpretation 'North Sweden' is inappropriate in the light of our distribution map. Type: not designated.
Fruit body annual, often persisting (as dead) till the following spring, made up of several layers of imbricate pilei, joined together by a basal plate, cream or chamois yellow when fresh, becoming brownish when dry. Total fruiting area up to 50 cm tall and 20-25 cm broad. Pileus up to 10 cm wide, projecting 3-6 cm from the substrate, 1-2 cm thick at the base,becoming thinner towards the margin. Upper surface rough. Margin sharp, inrolled when dry. Spines up to 12 mm long, cylindrical, acute, creamy yellow when fresh, becoming brownish and fragile when dry. Odour sour when fresh, becoming strong and unpleasant during drying, like rancid fat. Taste bitter.
Section: Context up to 15 mm thick, strongly fibrous and tough, sometimes with darker hygrophanous zones but not duplex, changing into trama without zonation.
Hyphal system monomitic. Generative hyphae 2-10 µm in diam, thin-walled, hyaline, variably clamped, branched, subparallel in the context, parallel in the trama. Septa of the context hyphae with single or multiple clamp connections; tramal hyphae clampless. Cystidia thick-walled, subulate or bottle-shaped, often apically encrusted. Basidia slender, 15-20 x 5-7 µm, without a basal clamp, four-spored. Spores ellipsoid, smooth, hyaline, 4.5-5 x 1.8-2.5 µm, IKI-, CB-.




 
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