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Description type:Non-original description 
Description:CLAVARIA COMPLANATA Cleland Trans. Roy. Soc. S. Australia 55: 159. 1931.
(ut C. complana).
Clavulina complanata (Clel.) Corner Ann. Bot. Mem. 1: 311. 1950. Holotype: ADW-15998, ?Hornsby, 13.vi. 1916, coil. J. B. Cleland; ?isotype-K, no. 68.
Single fruiting body 5.5 x 4.5 cm, repeatedly branched, apparently on soil with small twigs. Stem portion approximately 1.2 em long, composed of several flattened, connate stipes which branch irregularly upward; base
hispid-subtomentose, pale yellowish. Branches flattened, 3-7 mm thick, laterally expanded to 11 mm at the palmate arils, brittle, horny, subtranslucent in some places, and in those areas deep brick red or rusty red, but orange ochre in non-horny areas; internodes up to 1 cm long below, gradually diminishing to 6-7 mm apically ; arils lunate throughout; apices up to 2-3 cm long, about 1 mm thick, digitate to rounded.
Hyphae of lower portion of fruiting body uninflated, 4.5-10.5 µm diam, hyaline, somewhat thick-walled (wall up to 1.8 µm thick), strictly parallel, agglutinated, without clamp connections. Contextual hyphae of upper branches 2.4-5.7 µm diam, strictly parallel, hyaline, thin-walled, without, clamp connections. Subhymenium rudimentary; hyphae 1.6-3.4 µm diam, hyaline, thin-walled, tortuous to subparallel. Hymenium thickening; basidia 17.5-21 x.5.6-6.3 µm (hymenial measurements), clavate to subcylindrical, very` fragile, thin-walled, hyaline, easily collapsed, refringent under phase contrast when immature, without clamp connections, 4-sterigmate ; sterigmata up to 5.5 µm long, straight, spindly, somewhat divergent, almost always collapsed on basidia observed.
Spores 4.6-6.3(7.7) x 3.8-5.4 µm, globose, subglobose or broadly ellipsoid, smooth, hyaline, thin-walled, now aguttulate but with evidence of a single large guttule having been present; apiculus small, papillate, symmetrical; contents cyanophilous.
Fresh color: pale pinkish tussore, becoming brownish salmon; when damp staining paper pinkish salmon.
Corner (1950) described the genus Aphelaria for clavarioid fungi with tough consistency, flattened branches and hyaline spores. Later (Corner, 1953) he divided the genus into the subgenera Aphelaria and Tremelloden dropsis, the latter being elevated to genus rank by Crawford (1954). Corner (1966b) took issue with Crawford's disposition of this group of species, but retained the two genera derived by her.
The label of the type specimen bears notes by Cleland (and Cleland, 1931) indicating his recognition of, and separation from Wakefield's (1922) species Clavaria flabellata [-Aphelaria flabellata (Wakefield) Corner]. Both Wakefield (1922) and Corner (1950) described that species as white, while Cleland's notes state the fresh color of the type of C. complanata to be "pinkish tussore." On this basis the two may be separated, and a new combination in Aphelaria must be made. I propose Aphelaria complanata (Cleland) Petersen, comb. nov. (Basionym: Clavaria complanata Cleland).
Corner (1950) disposed of C. complanata under the genus Clavulina, stating, "Though its basidia are unknown, I have no doubt this is a Clavulina allied, perhaps, with C. cartilaginea." This conclusion was erroneous, especially in view of the morphology of the basidium as crucial to the taxonomy of Aphelaria as well as Clavulina.
Cleland's originally stated name, C. complana, was changed by him (1935), and the changed name adopted by Corner (1950). The changed name is obviously orthographically correct.
 
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