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Page number:163 
Description type:Non-original description 
Description:Multiclavula affin. coronilla (Martin) Petersen, Amer. Midl. Nat. 77:213-1967. = Clavaria coronilla Martin, Lilloa 5:194. 1940.
Fruitbodies 4-24 x 2-3mm, simple clubs or occasionally irregularly furcate above, arising from a small basal pad with embedded algae, narrowly fusiform to narrowly subclavate, gregarious to scattered, lemon yellow at base, off-white above ("pale cinnamon pink," "cartridge buff"); consistency tough, leathery; apex usually tapering to a point, occasionally minutely rounded. Odor and taste negligible.
Tramal hyphae 2-6 µm diam, hyaline, thin- to thick-walled (wall up to 0.2 µm thick), agglutinated, obscurely clamped, occasionally inflated abruptly, generally parallel; agglutinating substance apparently soluble in 3% KOH, granular. Hµm enium thickening; basidia 22-28 x 5-7 µm, suburniform, often geniculate, clamped; contents increasingly multiguttulate through development; sterigmata (4-)5-6, very slender, sublyriform.
Spores 5.0-6.1 x 2.9-3.2 µm (E = 1.67-2.13; Em = 1.89; Lm = 5.72 µm), cylindrical to suballantoid, flattened adaxially, thin-walled, hyaline, smooth; contents 1-severalguttulate, the guttules refringent and often masking the outline of spore wall; hilar appendix small, papillate.
Commentary: Corner (1950) summarized the taxonomic difficulties in the group, which exhibits few concrete characters on which to base identification. Petersen (1967a) relied on habitat (wood vs. soil; sµm biont type), fruitbody color (white vs. pigmented), num ber of sterigmata per basidium (four vs. extranumerary), and spore statistics on which to base taxonomic separation. Petersen (1967b) redescribed the type specimen, reiterating Martin's description of wood as habitat. Later (Petersen, 1967a), soil was added as a habitat, and even recently (Petersen, 1988), soil was allowed as a major habitat for the species, perhaps violating Martin's original concept.
The sole Chinese collection conforms to all characters stated by Martin, including: 1) wood habitat; 2) whitish fruitbodies; 3) extranumerary sterigmata (although not 6-8 as Martin reported and Petersen confirmed from the type specimen); and 4) appropriate spore dimensions. In fact, the specimen exactly fits the description by Thind (1961) of a northern Indian specimen on Piceawood. At Chang Bai Shan, the fungus was found in mixed mesophytic forest, on a large log, probably of a broadleafed tree (although possibly Pinus).
The specimen was field-identified as M. mucida, a worldwide taxon on alga-covered wood in temperate forests. From M. mucida it differs in extranumerary sterigmata. Thind (1961) used his specimens to conjecture that M. mucida and M. coronilla might be part of a taxonomic cluster, which only reinforces Corner's (1950) comment on taxonomic difficulties in the group.
Because the separating character, extranumerary sterigmata, is difficult to observe, little faith can be placed in reports of M. mucida. I have reported on an organism I called M. coronilla from high altitude in the southern Appalachian Mountains and from New Zealand, but the habitat of these collections was alga-covered soil or rock, and therefore included different algae from wood-inhabitors.
Martin (1940) drew attention to the urniform basidial morphology of M. (Clavaria) mucida. Hubbard and Petersen (1979) examined basidial nuclear behavior in that species, and showed it to be stichic, identical with that found in corticioid fungi with similar basidial development and morphology.
Specimen examined: Jilin Province, Antu County, Chang Bai Shan, on ? hardwood log, 18.VIII.87, coll. RHP, no. 47467 (TENN).
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