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Page number:222 
Description type:Non-original description 
Description:Ramaria versatilis var. violaceibrunnea (MARR & STUNTZ) PETERSEN comb. nov. - Figs. 9, 12
Ramaria fennica var. violaceibrunnea MARR & STUNTZ (1973). Biblioth. Mycol. 38: 78.
Ramaria violaceibrunnea (MARR & STUNTZ) PETERSEN (1986). Acta Bot. Yunnan. 8: 293.
Type (holotype). - USA; Washington, Mason Co., Mason Lake; 1. XI. 1966, coil. MARR 512 [n. v.], SUCO.
Illustrations. -MARR & STUNTZ (1973: pl. 4, figs. 21, 22, pl. 20, fig. 80).
Fruitbodies up to 13 x 6 cm, narrowly ellipsoid, truncateobovate to sphaeropedunculate in outline. - Stipe up to 40 x 30 mm, single, rounded at base, mycelial to rooting with many ropy, white rhizomorphs, smooth, not glabrous, brunnescent in creases (but not over surface or upon bruising), solid, upward pallid greyviolet ("vinaceous drab," "dark vinaceous brown," "light grayish vinaceous"); flesh white, solid, moist but not gelatinous or slippery, homogeneous, drying punky, somewhat friable. - Major branches 2-4, more or less terete, concolorous to branches above. Branches in 3-5 ranks, terete, avellaneous to purple in youth ("anthracene purple," "avellaneous," "drab"), darkening with spore production to smoky olivaceous shades ("buffy brown," "deep olive buff," "light brownish olive"); axils narrowly rounded; internodes diminishing gradually at maturity. - Apices double-dichotomous in youth, lengthening by maturity to digitate, buffy olive ("buffy olive," "light brownish olive"), darkening age ("dark olive buff," "citrine drab," "buffy brown"). - Odor sometimes weakly fragrant, taste negligible.
Macrochemical reactions. - FSW, FSA, FCL = strongly positive; KOH = bright peach red ("grenadine," fading to "rufous"); SYR = capricious; IKI, CRE, ANO, ANW, PYR, PHN, GUA = negative.
Tramal hyphae of stipe 4-10 µm diam, hyaline, thin- to thick-walled (wall up to 0.5 [an thick), conspicuously clamped, free, interwoven; ampulliform swellings at clamps, up to 14 µm broad, thin- to thick-walled (wall up to 0.5 µm thick), unornamented to scarcely and delicately ornamented; gloeoplerous hyphae not observed. - Tramal hyphae of upper branches 4-10 µm diam., hyaline, thin-walled, conspicuously clamped, free, interwoven; ampulliform swellings and gloeplerous hyphae not observed. - Subhymenium rudimentary, densely interwoven. - Hymenium thickening. - Basidia 50-60 x 7-9 µm, clavate, clamped; contents homogeneous; weakly cyanophilous; sterigmata 4, straight, spindly.
Spores (Fig. 9) 9-13 x 4.3-5.4 µm (E = 1.8-2.57; Em = 2.13; Lm = 10.42 µm), ellipsoid to ovate, flattened adaxially, obscurely roughened in profile; contents homogeneous to obscurely uniguttu late, the guttules dark, hardly refringent; wall up to 0.2 µm thick; hilar appendix eccentric, broad, with no throat; ornamentation of many small, isolated cyanophilous warts up to 0.1 µm high, without discernable orientation.
Commentary. - MARR (in MARR & STUNTZ, 1973) furnished a detailed description of R. fennica var. violaceibrunnea, to which my observations agree totally. MARR and I collected the holotype to gether, and an isotype is at TENN. The TENN portion was not properly dried, and the SUCO portion is much better preserved. Recently, MARR (1985, pers. comm.) suggested that there was a cline from typical cinnamon brown coloration of mature fruitbodies of R. violaceibrunnea to a somewhat olivaceous cast reminiscent of R. fennica. In western collections I also noticed this tendency, with west coast fruitbodies producing spores "deep olive buff" to "isabella color" in prints, unlike the more cinnamon colors of the typical variety. All of the fruitbodies gathered showed purple to lilac branches, however, not the colors and stature of R. fennica from Europe or eastern North America. Indeed, I have not seen fruitbodies of R. fennica in western North America. Only R. versatilis (vars. violaceibrunnea and versatilis) seem to represent the complex there.
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