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Page number:187 
Description type:Non-original description 
Description:Echinochaete russiceps (Berkeley & Broome) Reid, Kew Bull. 17: 285. 1963. Figs. 6a-d.
Polyporus russiceps Berk. & Br., Jour. Linn. Soc. Bot. 14: 48. 1873.
Polyporus grammocephalus var. russiceps (Berk. & Br.) Cooke, Grevillea 13: 84. 1885.
Leucoporus russiceps (Berk. & Br.) Pat., Bull. Soc. Mycol. France 30: 40. 1914.
Asterochaete russiceps (Berk. & Br.) Bond. & Sing., Ann. Mycol. 39: 58. 1941.
Dendrochaete russiceps (Berk. & Br.) G. H. Cunn., N. Z. Dep. Sci. Industr. Res. Bull. 164, p. 103. 1965.
Sporophores 2.3-6.0 cm wide, 2.5-7.0 cm measured radially, spathulate, narrowed behind to a broad, flattened, stipe-like base, 5-10 mm wide. The surface is now uniformly reddish brown with an ochraceous tint, and minutely scurfy tomentose under a lens, but there are also a number of dark radiating lines, from each of which arises a dark upstanding scale. A coloured drawing of the fresh plant shows it to have been radially streaky and reddish brown with scattered dark brown scales which appear much more prominent than in the exsiccatum. The margin of the pileus is also shown as ciliate, although there is no evidence of this in the dried material. Pores about 3-4 per mm, angular, with thin, papery dissepiments, and more or less isodiametric. In the fresh plant they are whitish, but they dry to an ochraceous buff. Tubes up to l.5 mm deep. Flesh up to 5 mm thick behind, but becoming very thin at the acute margin, whitish when fresh but ochraceous buff in dried material, and of very light weight. Hyphal structure dimitic, with generative and bovista-type binding hyphae which are distinctly dextrinoid. In the dissepiments many of the binding hyphae are more compact and have reduced spine-like branches. "Setae" present in the hymenium, often in groups, but very scanty. They are up to 30 µm long, and 8 µm wide, lanceolate, with a number of projecting spines near the apex and with a distinct dark brown wall. There are similar `setoid' structures on the surface of the pileus, where they are extremely abundant, but much larger, altogether more lax, and appear as dark brown, setoid, spinous hyphae. Spores mostly collapsed and therefore the measurements may not be entirely reliable, but 8.0-9.0 x 3.75-4.75 µm, thinwalled, hyaline, non-amyloid, and elongate-elliptic.
The above description is drawn exclusively from the type material: Sri Lanka, Peradeniya, Nov. 1868, No. 196. Other specimens from Sri Lanka which are definitely conspecific with it, i.e. Ancoombra, coil. T. Petch, Dec. 1917; Gangaruwa, coil. T. Petch, Aug. 1919-have better preserved spores with a range of 9.0-11.2 x 3.2-4.2 µm, which appear almost subcylindric. There also seems to be some variation with regard to the setoid elements on the cap, for while they are typically brown in colour and therefore very obvious in some collections, in other gatherings these elements remain more or less hyaline and inconspicuous. This may be a question of age of the fruitbody.
Echinochaete russiceps appears to be widespread in the Indo-Pacific. There are collections at K from: ASIA: Sri Lanka: Peradeniya, Nov. 1868, No. 961 [Type of P. russiceps); Ancoombra, coil. T. Petch, Dec. 1917; Gangaruwa, coil. T. Petch, Aug. 1919. Malaya: Forest Reserve Lompong, Johore State, coil. M JaquenoudSteinlin, 30 July 1974. Sarawak: near Long Kapa, Mount Dulit (Ulu Tinjar), 2500 ft alt., coil. P. Synge (No. 343), 27 Aug. 1932. Philippines: Forest Nursery, Quezon City, South Luzon, coil. Uyenco (No. 129), 17 June 1961. AUSTRALASIA: New Caledonia: on Casuarina cunninghamiana Miq., River Thi, coil. B. Huguenin (R. 256), 21 March 1965. New Guinea: Taun Creek, Bulolo, Morobe District, 3000 ft alt., coil. P. A. Wright (FM 33), 10 Nov. 1967. New Zealand: Tararuas, coil. F. Duguid (No. 162), 25 May 1947; on Beilschmiedia tairiri Benth. & Hook. f., Auckland, Kawau Island, coil. J. D. Atkinson, Dec. 1947.
It should be noted that some of these collections may ultimately have to be reassigned to other species, but until further material is available they are best left here since they appear to intergrade.
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