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Page number:308 
Description type:Non-original description 
Description:GANODERMA RESINACEUM (Boud.)Pat., Bull. Soc. Mycol. Fr. 5: 72. 1889.
=G. chaffangeoni Pat., ibid. 5: 74. 1889 (Fide Steyaert). = F. resinaceus (Boud) Sacc., Syll. Fung. 9: 179. 1891. = G. sessile Murr., Bull. Torrey bot. C1. 29: 604. 1902; North Amer. F1. 9: 120. 1908 (NY!). = G. polychromum (Copel.) Murr., North Amer. Fl. 9: 119. 1908. = G. pulverulentum Murr., ibid. 9: 121. 1908 (NY:). = G. praelongum Murr., ibid. 9: 121. 1908 (NY!). = G. subincrustatwn Murr., ibid. 9: 120. 1908 (NY!). = G. argillaceum Murr., ibid. 9: 122. 1908 (NY.). = G. nitidwn Murr., ibid. 9: 123. 1908 (NY!). = G. subperforatum Atk., Bot. Gaz. 46: 337. 1908. = G. platense Speg., Bol. Acad. Nac. Cienc. Cordoba 28: 363. 1926 (LPS:).
Figs. 7-10; 26-33; 60,
Annual, lignicolous, generally dimidiate to reniform, sometimes circular, spathulate or ungulate; isolated or imbricate, sometimes several pilei laterally confluent with their pseudostipes free or fused into one; sessile, substipitate; 3,5-55 x 3-18 x 1-8 cm. Pileus surface applanate, concave or more or less infundibuliform, smooth, irregularly rugose, concentrically sulcate and radially rugose or strong ly tuberculose, laccate, brilliant or dull, sometimes dull due to a thick deposit of spores. Central zone yellowish brown, very light in young specimens (Pl. 12 H 12), darkening with age from the centre towards the margin, with a broad cream coloured to yellowish marginal band (Pl. 9 E 4); in mature specimens there is no such gradation, the surface being dark reddish brown (Pl. 7 L 11) or light reddish brown, homogeneous (Figs. 8, 10). Margin sterile, thick, blunt, straight, incurved or recurved, yellowish cream in actively growing specimens, dark reddish brown in mature ones. Pseudo stipes lateral or central, very short to long, slender to thick, black, laccate, brilliant, 4-7 cm long, 2-5 cm wide, sometimes rudimentary. Section thin to very thick, thickening towards the base, 1-4 cm at about half the radius. Context corky and soft or woody and hard; 2-5 cm thick, uniformly brown (P1. 13 H 9), or with a thin darker band above the tubes. Hymenophore poroid, white to yellowish, becoming dark brown when old (Figs. 7, 9) with a layer of tubes often decurrent on the stem, 5-15 mm long, slightly lighter than the context; pores circular, large to medium sized, 2-5 per mm, 89-309 µm diam., dissepiments 27-267 µm wide. Hymenium composed only of basidia that are globose, soon collapsing, 714 x 8-20 µm, 4-spored (Fig. 60). Basidiospores of the "smooth" type, 9-13 x 5-8 µm, with a thick endosporium, light yellowish, ellipsoid, with numerous slender endosporic pillars that do not influence the perisporium, which is smooth and thin, thus appearing "smooth" when observed with the 0. M. at 400 x (Figs. 26-33). Dermis of the "hymenodermis" type, composed of claviform elements with scant lumen and blunt ends, originating from skeletal hyphae and arranged as in a hymenium, 7-16 µm diam., dermis 13-43 µm thick (Fig. 62). Hyphal system trimitic, with clamped, thin-walled generatives, with septa restricted to clamps, 1-6 µm diam., sparsely branched, abundant at the growth margin of the pileus and dissepiments (Fig. 71), Skeletals of the "arboriform type, clampless, aseptate, with a thick, golden wall, with few branches limited to the distal end, 3-8 µm diam., very long, forming the bulk of the context and dissepiments (Fig. 68). Binding hyphae of the "Bovista" type, clampless, aseptate, of limited growth, thick-walled, in general thinner and paler than the skeletals, much branched, 1-4 Aim diam. (Figs. 69-70), only present in the context, as a rule very scarce.
Hosts: on dead trunks of Tipuana tipu (dead as a result of the fungal attack); Quercus suber and undetermined hardwoods; at the base of Casuarina cunninghamiana, Platanus acerifolia, Ulmus procera, Acacia sp., Salix, Prosopis aLgarrobilla, Blepharocalyx tmeedii and Robinia pseudoacacia; also on stumps and roots of undetermined hardwoods. Domanski et al. (1967) record it also on Fagus and Alnus, although rarely.
Distribution: SOUTH AMERICA: Argentina (Buenos Aires, Capital Federal, Cordoba, Corrientes, Misiones, Tucumin and Salta); Uruguay; Venezuela. NORTH AMERICA: U. S. (Connecticut to Missouri, Alabama, Louisiana). CENTRAL AMERICA: Cuba, Honduras, Jamaica. It has also been recorded from Euro-Asia and Africa (Central). It seems to have a world-wide distribution in temperate and tropical areas.
Holotype: Fomes resinaceus Boud. (PC!).
Steyaert (1972) studied the holotypes of G. sessile, G. praelongum, G. argillaceum, G. resinaceum, G. polychromum, G. subperforatum and G. chaffangeonii, and found that they all possessed identical micromorphological features, particularly the "smooth" type of spores, for which reason he cons idered them synonyms, the valid name of the species being Ganoderma resinaceum (Bond.) Pat. We have been able to study all the holotypes of Murrill's species deposited at NY and we agree with him, but found that to the above list must be added G. nitidum and G. pulverulentum, as well as G. platense Speg., the holotype of which was studied in detail (LPS).
The description of G. subincrustatum given in North American Flora by Murrill, does not agree with the holotype (Murrill, 1908: 122), since the spores are given as 8 x 4 ?, whereas our measurements are 9-11 x 6-8 ?. In the diagnosis of G. nitidum Murr., in the same paper, no mention is made of the spores; its holotype, according to our measurements, has spores measuring 9-13 x 5-7 ?.
Spegazzini (1926) reported from Argentina G. lorentzianum Kalchbr., including stemless, pleuropodal and centrally stipitate specimens, a concave, more or less infundibuliform pileus, and ovate, smooth to slightly ornamented spores measuring 10-12 x 5-7 µm; at first we believed they were mere forms of G. resinaceum, but a detailed study of them showed there were really significative differences between those specimebs and G. resinaceum. We do not know whether Spegazzini studied authentic materials of G. lorentzianum Kalchbr.,and we were unable to locate the holotype in order to solve the problem.On the other hand of two other specimens identified as G. lorentzianum by Spegazzini at LPS, one has "smooth" type of spores and the other "semirugose" type. Thus, the former must be included in G. resinaceum and the latter in G. tuberculosum.
The study of the holotype of G. lorentzianum Kalchbr. would be important to ellucidate the cospecificity of this species with G. resinaceum, since should they prove to be the same, G. lorentzianum would be the valid name.
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