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Page number:491 
Remarks (public):The Phellinus igniarius complex is probably the most difficult complex of the whole genus. Numerous species have been described on basis of host specificity. The microscopical characters used to separate these species are in the best of cases marginal and are in general difficult to use. Studies have shown that there is partial incompatibility between these "species", but as long as there are no distinct morphological differences between them, we have adopted a wide circumscription here (like that of Niemelä 1975). It may be that we here are confronted with what has been called sibling species (see Hallenberg 1987) where genetic barriers have been established without distinct morphological differentiation.
Niemelä (1972, 1974, 1975, 1977) and Fischer (1987) should be consulted for a detailed discussion of the P. igniarius complex.
Description type:Non-original description 
Description:Phellinus igniarius (L.:Fr.) Quél., Ench. Fung., p. 172, 1886. - Boletus igniarius L. Spec. Plant. p. 1176, 1753. - Polyporus igniarius L.:Fr., Syst. Mycol. 1:375, 1821. - Fomes trivialis Bres., Icon. Mycol. 20:995, 1931. - P. igniarius var. cinereus Niemelä, Ann. Bot. Fenn. 12:110, 1975. - Ochroporus cinereus (Niemelä) Fischer, Bibl. Mycol. 105:159, 1986. - Ochroporus ossatus Fischer, Bibl. Mycol. 105:159, 1986. - Phellinus ossatus Fischer, Bibl. Mycol. 107:37, 1987, superfluous and illegitimate name, based on same type as O. ossatus. For comments on the nomenclature and typification of this epithet, see Parmasto 1988.
Basidiocarps perennial, sessile or rarely effused-reflexed, ungulate or sometimes
applanate, up to 11 x 20 x 8 cm; upper surface gray or blackish, glabrous, sulcate, becoming deeply rimose, encrusted; margin concolorous and glabrous or yellowish brown; pore surface pale cinnamon brown to dark purplish brown, the pores circular, 56 per mm, with thick, entire dissepiments; context dark reddish brown, zonate, woody, up to 2 cm thick; core absent or present next to substratum, with white tissue intermixed; tube layers concolorous with context, the tubes white-stuffed, in distinct layers, each up to 4 mm thick.
Hyphal system dimitic; contextual hyphae of two types, some brown in KOH, thick-walled, distinct, with rare branching, aseptate, 2-5 µm in diam, some hyaline, thin-walled with occasional simple septa, very indistinct; tramal hyphae similar, 2-3 µm in diam.
Setae ventricose to subulate, abundant to rare, 14-17 x 4-6 µm; core setae present in some specimens, irregularly lobed and branched; thick-walled, up to 15 µm in diam. Basidia broadly clavate, 4-sterigmate, 9-13 x 6-7 µm, simple-septate at the base.
Basidiospores broadly ovoid to subglobose, hyaline, smooth, thick-walled, negative in Melzer's reagent, acyanophilous, 5-6.5 x 4.5-6 µm.
Type of rot. Uniform white rot of the heartwood of living and dead hardwoods. Cultural characteristics. See Campbell 1938; Niemelä 1975; Nobles 1948, 1958, 1965; Venal 1937; Stalpers 1978.
Sexuality. Heterothallic and bipolar (Fischer 1987).
Substrata. Living hardwoods of many genera, continuing decay in dead trees, most common on Betula then Salix and Alnus, also fairly common on Acer, Corylus, Malus, Prunus and Sorbus, more occasionally on Aesculus, Amelanchier, Carpinus, Carya, Castanea, Cratageus, Fraxinus, Hippophae, Hydrangea, Juglans, Laburnum, Morus, Populus, Pterocarya, Robinia, Pyrus, Syringa, Tilia and Ulmus. Once reported from Pinus. Since this species is defined both in a wide and a narrow sense, the lists of hosts should be interpreted with care. In Asia and North America also recorded on numerous other hosts.
Distribution. Throughout Europe to the North Cape area in Norway. Circumpolar in the boreal-temperate zone.
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