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Page number:671 
Remarks (public):Specimens with flattened aculei, which is the normal, are easily recognized, but the variation in shape of the aculei makes the determination sometimes difficult. Microscopically there are no really good characters. The thinwalled, acute gloeocystidia are too few to be really effective. Specimens with conical aculei could be mistaken for H. aspera, but the aculei are usually longer and the projecting hyphae somewhat different. The species has been mistaken for H. arguta but lagenocystidia are never present.
Whether or not the species as here described is the one E. Fries intended with his Hydnum spathulatum is not settled. We have not seen authentical material. There are some details in Fries's description that do not agree with H. spathulata in modern opinion. Fries describes e.g. the colour as "candidus" which never is the case with H. spathulata. As the species is extremely rare in Sweden it is unlikely that Fries ever saw it, as his collecting was evidently very restricted. However, the use of the name seems to be unanimous, and we select the Romell collection cited above as a neotype untill authentic material is traced.
Description type:Non-original description 
Description:Hyphodontia spathulata (Fr.) Parm., Consp. syst. cort. p. 123, 1968. - Hydnum spathulatum Fr., Syst. mycol. I p. 423, 1821.
Fruitbody resupinate, creamish to ochraceous, effuse, adnate, hymenophore odontioid with dense aculei of varying shape, mostly flattened, but may be conical or subcylindrical, incised or denticulate, apically fimbriate when intact., about 1-2 mm long, on vertical substrate sometimes longer, margin variable, often determinate with a whitish, narrow, sterile zone, fimbriate under the lens,
Hyphal system monomitic; hyphae distinct, somewhat thichwalled, 2,5--3,5 µm wide, richly branched, irregularly interwoven in the subiculum, parallelly arranged in the aculeal trama, cyanophilous; projecting hyphae in the aculeal apices generally tapering, with attached aggregated crystals.
Cystidial organs of two kinds,-1) sterile, capitate hyphal ends, 25-50 µm long to the first septum, with apical resinous encrustation, frequently occurring in the hymenium,-2) thinwalled, acute cystidia (gloeocystidia) with one or more constrictions and dense, strongly stainable protoplasm, 20-35 µm long, sparsely occurring in the hymenium.
Basidia at first subclavate, then subcylindrical with suburniform constriction, 12-20 x 4-5 µm, with 4 sterigmata and basal clamp.
Spores ellipsoid, smooth, thinwalled, 1-3-guttulate, 4,5-5,5 x 3,5-4 µm.
Habitat. On decayed wood preferably of conifers but also occurring on deciduous wood.
Distribution. In Scandinavia found only once (Uppland: Tierp par., Prästön in river Dalälven, on deciduous wood, 1913-08-17, L. Romell 4981, Hydnum subfurcatum Romell ad int.) and it must be extremely rare. It seems to be rather frequent in E. Central Europe, e.g. in the Carpathians in Poland and Czechoslovakia, where it is one of the most frequent species of the genus. Parmasto reports it from Soviet and it occurs in N. America as well.
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