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Description type:Original description 
Description:Ramaria eryuanensis Petersen et Zang,, sp. nov. Figs. 1 , 2
Fruitbodies (Fig. 1 ) up to 12 x 8 cm, obovate in outline. Stipe up to 7 x 5 cm, massive, rounded at base, sometimes rooting, smooth, somewhat flattened, often with a few abortive stumps high on stipe, somewhat flattened laterally, white at base, sometime suffused pinkish at base, off-white above, easily brunnescent on all surfaces where handled or rubbed; flesh white, solid, not gelatinous or slippery, sometimes slowly sordid grey where cut. Major branches 2-4, stout, short, more or less terete, off-white ("pale ochraceous buff, "pale pinkish cinnamon"). Branches in 3-5 ranks, short, elongating reluctantly, crowded, more or less terete, pallid rosy beige ( "pinkish buff" ), easily brunnescent ("Verona brown"); flesh off-white ("pale cinnamon pink"), fibrous; internodes short, hardly diminishing above; axils narrowly rounded throughout. Apices digitate to awl-shaped, long, crowded, pastel rosy tan ("light pinkish cinnamon," "light ochraceous cinnamon"), easily brunnescent. Odor faintly fragrant; taste negligible to faintly fabaceous.
Macrochemical reactions; SYR = slowly positive, sometimes on rind only; IKI, FCL = positive; ANO, GUA, PHN, PYR, KOH, NOH, CRE = negative to equivocal, ANW = negative.
Stipe tramal hyphae 3-11 µm diam, hyaline, thick-walled (wall up to 1 µm thick), clamped, gnarled, freely branched, interwoven; ampulliform inflations as clamps or asymmetrical lobes, not unusually thick-walled, with extensive stalactitiform ornamentation; gloeoplerous hyphae not observed. Tramal hypliae of upper branches 2-8 µm diam, hyalie, thick-walled (wall up to 1 µm thick), straight, occasionally branched, parallel, not agglutinated or adherent; ampulliform clamps up to 15 µm broad, not unusually thick-walled, asymmetrical, unornamented; gloeoplerous hyphae not observed. Subhymenium extensive; hyphae 2-3 µm diam, very tightly interwoven, clamped. Hymenium thickening; basidia 68-75 x 9 -14 µm, clavate, clamped; contents densely granular; sterigmata 4, stout, straight.
Spores (Fig. 2 )10.8-13.0 x 4.0-5.0 µm (E = 2.31-3.00; Em = 2.65; Lm = 11.88 µm ), subboletoid to narrowly ellipsoid, smooth in profile; contents homogeneous and subrefringent or with 1-several refringent, ill-defined inclusions; wall up to 0.21 µm thick; hilar appendix prominent, with a throat; ornamentation of coarse striae, arranged in abaxial-distal to adaxial-proximal orientation.
Commentary. Fruitbody stature is typical of subg. Ramaria, with massive stipe, abortive branchlets and short branching system. Because of this, supportive characters assume higher significance. In the subgenus, color of upper branches and apices, for example, is usually in red, purple or avellaneous shades, so the pallid cinnamon colors of the specimens cited below is somewhat unique. Only R. secunda (Berk. et Curt.) Corner sensu Petersen exhibits this coloration. Stipe flesh amyloidity has not been ascertained for that species, and spores are almost identical to those of the Chinese material (for an isotype at FH; 10.4-12.6 x 4.1-5.2 µm; Em =2.35; Lm =11.19 µm). But our concept of R. secunda includes fruitbodies with modest, short stipes and well-developed branch systems, unlike the Chinese specimens, and R.secunda fruitbodies are not noticeably brunnescent.
Berkeley described fruitbodies of Clavaria secunda as "pale yellow," presumably on the advice of Curtis, who sent him the dried specimen. We have never seen such colors in this subgenus (except as stains on stipe surfaces), but in all other characteristics, the FH isotype specimen is matched by collections of a taxon from the southern Appalachian Mountains, leading to my concept of the species. Only the pallid cinnamon color differs from the type description and specimen. Thus I must cite the taxon as sensu Petersen, for perhaps pale yellow fruitbodies will yet be found.
Wu (1986) may have treated this as R. rubrievanescens, especially in light of the pallid pinkish shades which sometimes suffused the stipe base. She gave spore dimensions as 10--12.3 x 3-5.4 µm. Petersen has found no sign of pink coloration in the apices of R. eryuanensis, however, a diagnostic character for R. rubrievanescens,
Also quite similar is Marr and Stuntz's (1973) concept of R. strasseri. Stature and color is close, and spores are rather similar as well. Unfortunately, having examined several authentic specimens of R. strasseri, including co-types, We conclude that Marr and Stuntz's concept of that taxon was incorrect, as pointed out elsewhere (Petersen, 1986). If their concept of R. strasseri is contaxic with R. eryuanensis, it will have to be called by this name.
Fruitbodies of this species were among the most popular in the local markets of northern Yunnan. Fruitbodies of similar species are preferred for eating in western North America as well, especially in the spring. The Yunnan material reportedly fruited under Pinus.
Specimen TENN 47194 exhibited almost universally weakly positive macrochemical reaction, while the other specimens were inert except for IKI, SYR, and FCL.
Specimens examined; Yunnan, Lijiang Pref., Lijiang market, 30. viii. 86, no. 47306 (TENN, HKAS 20044); Lijiang marker, 2. ix. 86, no. 47299 (holotype, TENN; isotype, HKAS 20042); Er Yuan Pref., San Ying village market, 27. viii. 86, no. 47194 (TENN).
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