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Description:Stromata irregularly effused-pulvinate to elongate (a-c), 7-50 mm long × 3-22 mm broad × 1-1.4 mm thick, often growing beside and over old black depauperate stromata, with faintly exposed discoid perithecial contours (d); surface pruinose, Dark Vinaceous (82) to Brown Vinaceous (84), subsurface blackish, composed of weakly carbonaceous tissue and inconspicuous brownish black granules yielding dilute Olivaceous Buff (89) pigments in 10% KOH (e); subperithecial tissue 0.2-0.5 mm thick, fibrous, grey brown to blackish, seated on a thin black line (i). Perithecia lanceolate, 0.7-0.9 mm high × 0.3-0.35 mm diam, with perithecial contents Citrine Green (67) (f, i). Ostioles umbilicate, faintly papillate in old black stromata (d). Asci few, 90-110 µm total length (k), the spore-bearing parts 55-65 µm long × 4.5-5 µm broad, the stipes 30-45 µm long, originating in unilateral spicate arrangement from long ascogenous hyphae, with apical ring discoid, 0.8 µm high × 1.7-2 µm broad, bluing in Melzer’s reagent. Ascospores 7.5-9.4 × 3.2-4.2 µm (M=8.6 × 3.5 µm, n=30), ellipsoid equilateral with broadly rounded ends to oblong, pale brown (j), uniseriate in the ascus, with faint germ slit spore-length to slightly less than spore-length (h). Perispore indehiscent in 10% KOH; epispore smooth. Cultures and anamorph. Colonies (22 °C, 2 wk) on OA attaining 38-45 mm, at first velvety to felty, finally wooly to hispid, azonate, margins diffuse to fimbriate, pale yellow brown to greenish brown (4F6), finally dull grayish brown (5E3, 5F3), dull reddish gray (7E4-3, 7F4-3), occasionally with dark brownish black exudates in age, sporulation scattered on aerial hyphae after 2 to 3 wk, reverse dull yellowish gray (5E2, 5F2)to grayish red brown (8F3-2). On 2% MEA attaining 37-45 mm, felty, faintly zonate, margins fimbriate to submerged, pale gray to dull reddish gray (8E3-1, 8F3-1), finally dull gray brown, scant sporulation on the aerial hyphae, reverse deep reddish black (9F3-2) to black. On YM attaining 35-43 mm, appressed to felty, with radial hyphal strands running across the agar surface, pigmentation similar to MEA, scant sporulation on aerial hyphae, reverse with strong black mottling. On SNA attaining 30-40 mm, and 14 mm before agar desiccated, mostly submerged, hyaline at first but in age forming olive brown to grayish brown hyphal strand radiating from central inoculation point, with pale buff sporulation evident on agar surface, especially along axes of hyphal strands. Agar plugs of MUCL 49879 incubated at 37 °C on YM only developed scant hyphae from the agar plug, while MF5495 grew about 3-4 mm. Conidiophores on SNA arising from repent hyphae on agar surface, often flowing or concentrated along axes of hyphal strands, erect, up to 300 µm tall, 4-7 µm wide at the base, septate, hyaline to olive brown or grayish brown in 5% KOH, finely roughened, with scattered brownish gray pigment granules, slightly thick walled at the base, with up to 4 levels of branches, with first primary branch, between one- to two-thirds up the main axis, with 2-4, but usually 3, terminal conidiogenous cells, rarely with intercalary conidiogenesis. Conidiogenous cells usually arising from terminal cells, rarely with swollen intercalary loci, bearing a cluster of holoblastically produced conidia, determinate, cylindrical, mostly with a slightly clavate to inflated apex, finely roughened, bearing finely denticulate secession scars at and slightly below the apex, 10-22 µm × 2-4 µm. Conidia hyaline, buff to white in mass, smooth, ovoid to ellipsoid, with truncated basal scar, 4-7 µm × 2-3 µm, usually with a small guttule at each end.  
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