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Page number:1173 
Remarks (internal):Morchella snyderi corresponds to phylogenetic species Mel-12 in O'Donnell et al. (2011). A combination of ecological and morphological features separates the species: it appears in non-burned conifer forests in western North America; its stipe is often ridged and lacunose (although the stipes of many morels can become ridged and sublacunose, especially near the base, when mature, the stipe of M. snyderi is often prominently lacunose and ridged throughout most of its development) and prominently granulated; the colors of the hymenophore approximate those of esculenta-like morels when young, but the ridges become smoky brown to black with development or upon drying; its hymenophore is conical, with longitudinally arranged pits and a sinus at the point of attachment to the stipe; and its ascospores are comparatively large. Young specimens of M. snyderi can appear similar to M. frustrata, but the latter species has ridges that do not darken and has smaller ascospores. Morchella brunnea also is similar, but its hymenophore is more brown when young and its stipe is not regularly lacunose. We found one ascoma fragment in the putative holotype of M. crassistipa Snyder to be phylogenetically conspecific with M. snyderi, but we were unable to recognize Snyder's name because the type collection was mixed as to species (see supplementary materials, Type studies, for full discussion). Morchella snyderi was not treated in Kuo (2005). 
Description type:Original description 
Description:Morchella snyderi M. Kuo & Methven, sp. nov. Fig. 16
MycoBank MB 563959
Ascomata 60-140 mm high. Hymenophore 35-80 mm high; 30-50 mm wide at the widest point; conical; pitted and ridged, with 16-22 primary vertical ridges and frequent shorter, secondary vertical ridges, with occasional sunken, transecting horizontal ridges; attached to stipe with a sinus about 2-4 mm deep and 2-4 mm wide. Ridges glabrous or very finely tomentose; pale yellowish, becoming pale tan, then grayish brown with maturity and darkening to nearly black when dried; flattened when young but with age sometimes becoming sharpened or eroded. Pits more or less vertically elongated, at least at maturity; finely tomentose; yellowish when young, becoming pale tan to pale grayish brown. Stipe 35-70 mm high; 25-40 mm wide; more or less equal, or sometimes basally subclavate; at first finely mealy with whitish granules, becoming prominently granulated; whitish to pale brownish; usually becoming prominently ridged and/or lacunose with maturity. Context whitish; 1-2 mm thick in the hollow hymenophore, becoming layered and chambered, especially near the base of the stipe. Sterile inner surface whitish and pubescent. Ascospores 25-37 x 15-23 µm; elliptical; smooth; contents homogeneous. Asci 225-300 x 17.5-32.5 µm; eight-spored; cylindrical; hyaline in KOH (2%). Paraphyses 100-200 x 7.5-20 µm; cylindrical; apices rounded to subclavate, clavate, or occasionally subcapitate or widely fusiform; septate; hyaline to faintly brownish in KOH (2%). Elements on sterile ridges 75-175 x 10-20 µm; septate; terminal cell subclavate, clavate, subcapitate or widely fusiform; with hyaline to brownish or brown contents in KOH (2%).
Ecology. Appearing under non-burned, montane conifers, including Ps. menziesii, P. ponderosa and A. concolor; April, May and early June.
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