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 Add this item to the list  Trametes modesta (Fr.) Ryv.
Page number:123 
Description type:Non-original description 
Description:Trametes modesta (Fr.) Ryv.
Norw. J. Bot. 19 (1972) 236; Ryvarden and Johansen (1980) 576; Ryvarden, Mycotaxon 13 (1981) 182; Gilbertson and Ryvarden (1987) 750.
Daedalea modesta (Kunze) Aoshima, Trans. mycol. Soc. Japan 8 (1967) 2.
Coriolus clemensiae Murr. (1908), Ryvarden, Mycotaxon 23 (1985) 173. (See under Tr. badia, p. 72).
Pileus -3 cm in radius, 5 cm wide, sessile or shortly stipitate from a discoid base, often imbricate, dimidiate, spathulato-flabelliform when stipitate, rarely shortly effuso-reflexed, matt, dull, radially subrugulose, inoderm, often developing a thin pad at the base, dull cinnamon rufous or warm brown with narrow lighter zones, often paler at the base and somewhat fuscous; margin subacute, pallid tan (not white). Stem 2-5 x 3-5 mm (when present) with a discoid base 4-10 mm wide. Tubes 0.5-0.8 mm long, less than 0.5 mm at 5 mm from the margin of the pileus, very short, pallid dingy tan; pores 85-105 µm wide, dissepiments 45-100 µm thick, round or elliptic, entire, pallid dingy tan. Flesh 1-2 mm thick at the base of the pileus, 510 mm at the extreme base, 0.5 mm at 5 mm from the margin, corky floccose, pale fawn tan. Smell sour, not strong.
On dead wood in the forest. Pantropical.
Spores 4 x 2 µm, white, smooth, subcylindric, inamyloid; 4.5-6 x 1.5-2 (-2.5) µm, Ryvarden and Johansen. Basidia 10-16 x 4.5-6 µm, sterigmata 4, 2.5 µm long. Hymenium compact, not thickening. Cystidia and hyphal pegs none. Hyphae as in Tr. feei but: 3-4.5(-5) µm wide in the flesh, 2.5-3.5(-4) µm in the dissepiments; binding hyphae not so numerous, scattered throughout the flesh but very abundant in the lower layer 100-150 µm thick over the tubes, rather scarce in the dissepiments (composed mainly of skeletals), never concealing the structure of the palisade on the pileus. Surface of pileus with a close palisade of skeletal ends, not dilated, but becoming progressively inclined over the limb and inoderm in its outer part.
Collections: Malaya, Negri Sembilan, Angsi Forest c. 250 m alt., 4 July 1930, Sing. F.N. 23715; Sing. F.N. 11619, leg. Md Nur 1923.
The description which I give is based on these two collections. Sing. F.N. 11619 was identified by Lloyd as Tr. modesta, but both collections were referred to Tr. confundens Ces. by E.M. Wakefield with the note that they had darker flesh and more silky pileus than Tr. modesta. Be that as it may, the description of Tr. modesta by Ryvarden and Johansen indicates a similar fungus with minute pores and variable colour from pinkish brown through brown to pallid tan. They give the spores as slightly longer and, even more so, in the neotropics 6.5-8 x 3-3.5 µm (Ryvarden 1981). The fungus tends in some respects to Tr. feei and in others to Tr. biogilva.
However, Gilbertson and Ryvarden separate Tr. feei in Fomitopsis on account of its brown rot and retain Tr. modesta with a white rot.
I think that there are several species or varieties about Tr. modesta which can be settled only by accurate description of the living, accompanied by ascertainment of spores and hyphal construction. I have listed such as I have found as var. A, B and C, with the indication to compare the varieties of Tr. biogilva. In these varieties, the ends of the skeletals in the palisade on the pileus become slightly encrusted with brown matter, which causes their deeper or richer colour, as in Tr. fulvirubidus. The cinnamon tints seem to be caused by a mixture of pinkish and pale brown in the hyphal walls.

Key to the varieties of Tr. modesta
1. Pores 80-120 µm wide.
2. Pileus warm brown to cinnamon rufous, sessile to substipitate with discoid base. Flesh 1-2 mm. Malaya. T. modesta
2. Pileus fawn brown, sessile. Flesh 4-5 mm thick. Palisade on pileus thinly encrusted. Borneo var. A
1. Pores wider. Palisade on pileus thinly encrusted.
3. Pores 130-200 µm wide, white. Flesh pinkish. Malaya. var. C
3. Pores wider. Flesh brown.
4. Pores 200-500 µm wide, partly gyrose. Flesh -6 mm. Pileus inoderm, sessile or substipitate to a discoid base. Malaya, Borneo. Tr. biogilva (p. 78)
4. Pores 150-300 µm. Flesh 1-2 mm. Pileus sessile. Malaya. var. B

Trametes modesta var. A
Pileus -2.5 cm in radius, sessile, horizontal, flabelliform, imbricate, not sulcate, matt, fawn brown with slightly darker zones, paler to the margin. Tubes -2 mm long, dried concolorous with the flesh; pores 80-110(-120) µm wide, dissepiments 40-110 µm thick, entire to occasionally 2 confluent, pallid white. Flesh 4-5 mm thick at the base of the pileus, corky fibrillose, dried pinkish cartridge buff, no crust.
On a dead stump in montane forest. Borneo, mt Kinabalu, Mesilau 1700 m alt., 22 April 1964, RSNB 843 1.
Spores? Hymenium not formed. Cystidia and hyphal pegs none. Skeletal hyphae 4-6 µm wide, -7 µm in places, 6-8 µm in potash, walls -2 µm thick and pale pinkish ochraceous, unlimited or many ending in 1-3 stout binding hyphae 3-4 µm wide dividing into narrower branches with ultimate branchlets 1.5-2 µm wide, not flagelliform, produced also from the generative hyphae; dissepiments with skeletals 3-4 µm wide. Surface of pileus with a close palisade of thick-walled skeletal ends 100-150 µm long, the tips becoming embedded in a brown resinous exudate -15 µm thick, giving colourless oily masses in potash.
This fungus seems to lead to Tr. biogilvoides and to Tr. fulvirubida. It is possible that all the skeletal hyphae are more or less arborescent with terminal binding processes.

Trametes modesta var. B
Pilei -2 cm in radius, sessile, imbricate, slightly ascending, sometimes with a short resupinate foot, often connate laterally, thin applanate, rather coarsely inoderm in the outer part, scabrous fibrillose towards the base, pale fawn brown to subferruginous, faintly zoned; margin whitish. Tubes 2-3 mm long, c. 1 mm at 5 mm from the margin of the pileus, concolorous with the pileus; pores 150-300 µm wide, dissepiments 90-200 µm thick, angular, often slightly radially elongate, rarely 2-3 confluent, pallid white. Flesh 1-2 mm thick at the base of the pileus, sometimes with a basal pad of tissue -6 mm thick, corky fibrous, concolorous with the pileus, no crust.
On a fallen trunk in the forest. Malaya, Negri Sembilan, Bukit Senaling, 24 June 1930, Sing. F.N. 23714.
Spores? Cystidia and hyphal pegs none. Skeletal hyphae in the flesh 3-5 µm wide, 5-8 µm in potash, 2.5-4 µm in the dissepiments, with pale brownish glassy walls and more or less obliterated lumen; binding hyphae fairly abundant throughout the flesh, not flagelliform; generative hyphae abundant, clamped; dissepiments with the ends of skeletal and binding hyphae projecting to 20 µm into the tubes. Surface of pileus with a rather irregular and loose pile, in places fasciculate at the base of the pileus, directed obliquely forward over the limb and often in microscopic bundles, the skeletal tips often subclavate 4-6 µm wide, eventually thinly encrusted in places with yellow brown granular matter soluble in potash, not forming a distinct crust.
But for the wide pores, this could be squeezed into Tr. modesta, but compare Tr. multiflabellata (p. 126).

Trametes modesta var. C
As var. B, but the pores 130-220 µm wide, pallied white; flesh pinkish. Spores? Malaya. Negri Sembilan, Angsi Forest, 2 Dec. 1922, Sing. F.N. 9935, det. Polyporus didrichsenii by C.G. Lloyd; herb. Singapore.
The colour of the pores and flesh was noted on the field-label. P. didrichsenii is given as a synonym of Tr. menziesii by Ryvarden and Johansen, and the name cannot refer to this fungus.
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