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 Add this item to the list   Gloeoporus amorphus (Fr.) Clem. et Shear
   
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Page number:45 
Description type:Non-original description 
Description:Gloeoporus amorphus (Fr.) Clem. et Shear
Pilat, Atlas Champ. Europe ser. B. fasc. 13-14 (1937) 153; Teston, Bull. Soc. Natural. Oyonnax 7 (1953) 85.
Skeletocutis amorpha (Fr.) Kotl. et Pouz. Ceska Mykol. 12 (1958) 104; Ryvarden (1976) 415.
lt is nut clear to me that the construction of this common, north temperate, fungus has been adequately understood. lt is treated as having gelatinous flesh and tubes. but 1 do nut Eind this to be so in living material; certainly it is so in dried material restored by swelling and softening in potash. Then, the so called skeletal hyphae of the tomentum are the ends of thick-walled monomitic hyphae such as occur in many polypores with villous pileus. The occasional skeletal hyphae said to occur in the flesh are skeletal cells to be found in other species of the genus. as G. croceopallens. The living tissue of the flesh and tubes has an oily waxy excretion from the hyphae which gives the tissue its waxy translucence in the living state and its horny character when dried. The following are my notes on the structure of the living fruit-bodies.
Spores 3.5-5 x 1.3-1.5 µm. allantoid. I-ve. Tubes -1.5 mm long. waxy, nut gelatinous; pores drying yellow to orange. Flesh of the pileus composed of three layers; 1. the upper tomentum 1-1.5 mm thick. composed of thick-walled, mostly unbranched hyphae 3-5 µm wide, sparsely septate with clamps, rather loosely interwoven; 2. the middle coriaceous layer of waxy felted tissue 0.5-1 mm thick. nut gelatinous, composed of radiating clamped hyphae 3-5 µm wide, with thin or thickened walls, some with more or less linear lumen as intercalary skeletal cells; 3. the dense, almost solid, luwer layer c. 0.5 mm thick. immediately above the tubes. nut gelatinous, composed of radiating hyphae as in the middle layer and of binding hyphae 2-3 µm wide frequently branched and lobing with thickwalls, up to 100 µm.
long (? more) with subacute thin-walled tips, interweaving profusely, derived from the radiating hyphae of this layer (corresponding with the pore-field). Dissepiments composed of descending and interweaving narrow hyphae 1.5-3 µm wide, with thin or slightly thickened walls -0.7 µm, derived from the binding hyphae of the pore-field, neither gelatinous nor agglutinated. All hyphae, especially in the lowest layer of the flesh and in the dissepiments, with an oily waxy excretion, thick and slimy in places, becoming pale yellow to pale orange in the dissepiments (as the colour of the tubes and pores). Tissue of the middle and lower layers of the flesh becoming horny on drying, but on treatment with dilute potash more or less gelatinous with much oily matter; only the tomentum remaining non-gelatinous.
In the waxy texture of the flesh and tubes, G. amorphus seems to present the antecedent to the gelatinous texture more typical of Gloeoporus. In other respects the species closely resembles G. dichrous. I am puzzled, nevertheless, by a collection from India, determined by B.K. Bakshi as Polyporus amorphus Fr., namely Chakravati U. P. Mundali, leg. Bakshi 28 Oct. 1957, Forest Research Institute, Dehra Dun. This collection, without field-notes, and without certain spores, has apparently a truly dimitic construction in the flesh with unlimited and aseptate skeletal hyphae 3-5 µm wide. It reminds me of Gloeoporus sp.A from Japan (p. 62).
 
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