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 Add this item to the list  818668 Original description
Description type:Original description 
Description:Ascomata apothecial, collectively or solitary erumpent in early stage of development, emerging from longitudinal crack of the twig periderm, when fully expanded speciously superficial, at first globular, then hemispherically expanding, deep cupulate when mature, with only narrow basal area immersed in inner bark, sessile, ± circular from the top view, *0.4–1.3 mm and †0.3–0.7 mm diam, solitary or crowded (up to dozen apothecia). Hymenium steel grey, not wrinkled; margin sharp, whitish pubescent, entire, not lobed, permanently inrolled; excipular surface hazel brown, somewhat mealy rugulose. Basal hyphae macroscopically indistinguishable. Pseudostromatic hyphae not forming macroscopically evident demarcation lines. Asexual morph not seen. Hymenium *65–75 µm thick, consisting of asci and paraphyses. Asci cylindrical-ventricose with sub-conical apex, *68–79 × 8.5–10.5 µm, †55–61 × 5–6.5 µm, pars sporifera *28.5–34.5 µm, 8-(rarely 4-)spored, base cylindrical-truncate, arising from narrow repetitive croziers, apical apparatus moderately refractive in water, in Lugol’s solution apical ring strongly amyloid (3bb) of Calycina-type. Ascospores subscutuliform, bilaterally symmetrical, aseptate, *(10–)10.5–12.5–14(–14.5) × 2.5–2.9–3(–3.5) µm, †9.5–11.5 × 2–2.5 µm, *Q = (3–)3.5–4.3–5, †Q = 3.5–4.5, hyaline, smooth, sporoplasm containing several small to medium sized lipid bodies, bi- to triseriate when inside living asci, with abundant sheath enveloping whole spores when freshly ejected. Hamathecium composed only of paraphyses, cylindrical-lanceolate, more rarely clavate, apical cell *30–66.5 × 3–5(–5.5) µm, †20–53.5 × 3–4 µm, *containing several subhyaline, strongly refractive globose vacuolar bodies which readily coalesce in still living cells, usually abundantly exist as irregular remnants in water rehydrated material, KOH reaction negative, completely dissolving vacuolar bodies, wall thin and hyaline, sometimes covered with strongly refractive golden yellow patches which are not dissolved but discoloured in KOH. Subhymenium slightly differentiated from medullary excipulum, *5–7 µm thick at the upper flank, composed of hyaline *densely packed small ± cylindrical cells *2.5–4(–7) µm wide. Medullary excipulum devoid of crystals, moderately gelatinised (lilac in Brilliant Cresyl Blue), very reduced at the upper flank, composed of hyaline textura porrecta, *8–12 µm, †5–7 µm thick; considerably thicker at the middle flank, of textura intricata-porrecta *14–23 µm, †12–17 µm thick; cells *2.5–4.5 µm, †1.4–3.5 µm wide, inamyloid including the perihymenial area. Ectal excipulum altogether *32–37 µm, †31–34 µm thick at the upper flank excluding hairs, *52–68 µm, †46.1–51.8 µm thick at the middle flank, devoid of crystals, clearly differentiated into two layers: inner layer *27–36 µm thick at the middle flank, composed of hyaline textura angularis with cells *4.5–17 × 4–11 µm, cell walls refractive, 0.5 µm thick, intercellular spaces densely interspersed with minute irregular dull particles; outer layer *24–30 µm thick at the middle flank, composed of brown textura globulosa-angularis with cells *5–11 × 4.5–10 µm, cell walls ochraceous-yellow to tobacco-brownish, refractive, 0.6–0.9 µm thick, intercellular spaces with abundant dark-brown patches, outermost cells each containing a single olivaceous globose vacuolar body. Outermost cells on the upper flank giving rise 1–4-celled hairs, *(12.5–)20–60 µm, †10–46 µm long, running at high angle to the excipular surface, with ±moniliform cells *6–13.3 µm, †4.5–11.5 µm wide, wall tobacco-brownish, 0.5–1 µm thick; terminal cells broadest, clavate-rhomboid to shortly lanceolate, *12.5–28.5 × 8.5–13 µm, †9.5–18.5 × 8–12.5 µm, *containing single, olivaceous, highly refractive vacuolar body, wall brown to rusty brown, *0.5–1 µm, †0.5–1 µm thick, surface covered with dark brown thick patchy warts which are stable in KOH, texture without KOH coloured reaction, in Lugol’s solution completely inamyloid. Hairs on marginal rim flexuous, undulate, smooth, hyaline and thin-walled, cylindrical to subclavate, rapidly collapsing, *20–65 × 4–8 µm. Subiculum abundant, hyphae arising from lower and basal flank, wavy, almost not branched, evenly septate, with occasional short knotty lateral protuberations up to †5.5 × 4 µm diam., walls thickened, †0.5–1 µm, smooth, but darker brown hyphae adpressed to the excipular surface and some distant lighter coloured hyphae finely warted, hyphae mostly greyish yellow to hazel brown, †2.5–4 µm wide. Subperidermal structures: of thick-walled cylindrical hyphae similar to those in subiculum, but all finely warted, sparingly branched and without knotty protuberations, growing on surface and inside the inner bark tissue, †2–4 µm diam., walls †0.4–1 µm thick, compact stromatal structures absent. 
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