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 Add this item to the list  833375 Original description
Remarks (public):Ecology and distribution: Known only from the Galapagos; growing on sunny, wind- and rain-exposed lava rock, from the coast to the transition zone; easily overlooked due to its small size. Notes: With its black discs and creamy pale thallus L. ombligulata superficially resembles Tephromela, but its asci belong to the Lecanora-type and its epihymenium color also differs from the purple, N+red pigmentation observed in Tephromela. Because of its indistinct umbilicate growth, L. ombligulata closely resembles species of Rhizoplaca. It could be mistaken as a minute, very immature form of R. glaucophana (Nyl. ex Hasse) W.A. Weber. From that species L. ombligulata can, however, be distinguished by its unusual chemistry and the differently pigmented epihymenium (R. glaucophana contains only pseudoplacodiolic acid and it has a reddish brown epihymenium). Because thalli of L. ombligulata are so small, it is difficult to determine whether they are truly umbilicate; they may better be called peltate, with individual squamules attached at their center. Despite this growth closely resembling some thalli of Rhizoplaca, the conidia of L. ombligulata are short, bacilliform to baculiform, not filiform. The presence of xanthones in combination with diploicin is equally unusual, this secondary chemistry being unknown from species now placed in Rhizoplaca. Therefore, we decided to describe the new species not as a Rhizoplaca, but within Lecanora s.l. 
Description type:Original description 
Description:Thallus saxicolous, 0.1–3(–5.6) mm in diam., pale grayish olive, indistinctly umbilicate, of minute, scattered and widely dispersed ±peltate to ±cerebriform verrucae, closely adjoining verrucae often becoming conspicuously inflated, swollen and highly convoluted; surface dull to ±shiny, smooth, epruinose, but typically whitish eroded along the thallus folds; lacking soredia; prothallus absent. Apothecia sparse to numerous, occasionally crowded and slightly deformed by mutual pressure, circular or slightly undulate, (0.3–)0.4–1.2(–1.7) mm diam., soon becoming sessile and basally constricted, distinctly lecanorine with a thickened, entire margin, concolorous with the thallus, disc plane, with a highly roughened, finely pruinose surface; thalline exciple with cortex, photobiont layer and medulla, filled with abundant minute crystals, soluble in K (allophana-type); hypothecium not distinctly differentiated from subhymenium, pale brown, extending into the inner exciple; hymenium hyaline, not inspersed; epihymenium with a deep bluish green pigment (cinereorufa-green: intensifying in K, HCl+ bluish green, N+ reddish violet), lacking crystals; proper exciple thin, indistinct, with few small, granular crystals, insoluble in K; thalline exciple thick, distinctly corticate, abundantly filled with small crystals soluble in K, lacking large crystals; subhymenium and hypothecium hyaline, abundantly filled with small crystals, soluble in K; asci clavate, 8-spored, Lecanora-type; ascospores simple, hyaline, smooth, broadly ellipsoid to subglobose, rarely ±ovoid, thin-walled, (4.5–)7.5–10.1(–10.8) × (4.9–)5.4–7.1(–8.8) µm (n = 45). Pycnidia immersed, flask-shaped with a colorless wall, and deep bluish green ostiole (cinereorufa-green: intensifying in K, HCl+ bluish green, N+ reddish violet); conidia simple, elongate bacilliform ('baculiform'), (5.9–)6.8–9.8(–10.8) × 1.5–2 µm (n = 15). Chemistry: Thallus cortex P–, K+ yellow, C+ orange, KC+ orange, UV- deep orange; medulla P-, K-, KC+ orange, C+ orange, UV- deep orange; thiophanic acid [major], diploicin [minor], dechlorodiploicin [minor], isofulgidin [minor]. 
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